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More recently, another group has proposed an AMPK-dependent mechanism by which metformin reduces cAMP [33]: treatment of mouse hepatocytes with a more specific AMPK activator reduced glucagon-induced cAMP levels and this was traced to the direct AMPK-mediated phosphorylation of the cAMP-specific 3′,5′-cyclic phosphodiesterase 4B (PDE4B), triggering cAMP breakdown (Fig
